Keetia venosa (Oliv.) Bridson
  • Kew Bull. 41: 974 (1986) 


Notes: Distribution: Trop. Africa

General Description

Scandent shrub or climber 2–7 m tall; young branches sparsely to densely covered with rusty-coloured hairs.Leaf blades 4.5–14 × 2.5–6(8) cm, oblong-elliptic or narrowly to broadly elliptic rarely round, acuminate at apex, obtuse to rounded or rarely subcordate at base, glabrous or rarely glabrescent on upper surface, glabrous save for the sparsely to densely pubescent nerves, or occasionally sparsely pubescent beneath; lateral nerves in 5–9 main pairs; tertiary venation finely reticulate, always with element perpendicular to midrib the more conspicuous giving a scalariform appearance and frequently either raised or impressed above; domatia present as inconspicuous tufts of hair; petioles 5–15 mm long, pubescent; stipules 5–14(17) mm long, up to 5 mm wide at base, triangular at base, usually rather abruptly narrowing to a linear or acuminate apex, pubescent outside; leaves subtending lateral branches smaller, ± orbicular.Flowers 4–5(6)-merous, borne in 20–70-flowered pedunculate cymes; peduncles 5–17 mm long, pubescent or rarely glabrous; pedicels 2–5 mm long, pubescent or rarely glabrous; bracteoles 2.5–6 mm long, linear-lanceolate.Calyx tube 0.75–1 mm long, glabrous or pubescent; limb 0.75–1.25 mm long, divided into teeth usually for less than half its length, ciliate.Corolla creamy-white; tube 2.25–3 mm long, with a ring of deflexed hairs just below the throat inside; lobes 1.5–2.25 × 1–1.75 mm, lanceolate to ovate, acute.Anthers fully exserted but seldom reflexing.Style 4–6.5 mm long, glabrous; pollen presenter 0.75–1.25 mm long; disk pubescent.Fruit 8–11 × 11–15 mm, slightly indented at apex, glabrous, black when mature.Pyrene 6–7 × 5–6 mm, hemispherical to suborbicular, with lid-like area lying across the apex with a small crest.Seed with endosperm streaked with granules.

Scandent shrub or climber, 2–7 m. tall; young branches sparsely to densely covered with rusty coloured hairs. Leaf-blades oblong-elliptic or narrowly to broadly elliptic, rarely round, 4.5–14 cm. long, 1.5–6(–7.5–8) cm. wide, acuminate at apex, obtuse to rounded or rarely subcordate at base, glabrous or rarely glabrescent and shiny above, glabrous save for the sparsely to densely pubescent nerves or occasionally sparsely pubescent beneath, papery to subcoriaceous; lateral nerves in 5–9 main pairs; tertiary nerves finely reticulate always with element perpendicular to midrib more conspicuous and often raised or impressed above; domatia present as inconspicuous tufts of hair; petioles 0.5–1.5 cm. long, pubescent; stipules triangular at base rather abruptly narrowing to a linear lobe or acuminate apex, 0.5–1.4(–1.7) cm. long, up to 0.5 cm. wide at base, pubescent outside; leaves subtending lateral branches smaller, ± circular. Flowers 4–5(– 6)-merous, borne in pedunculate 20–70-flowered cymes; peduncles 0.5–1.7 cm. long, pubescent; pedicels 2–5 mm. long, pubescent; bracteoles linear-lanceolate, 2.5–6 mm. long. Calyx-tube 0.75–1 mm. long, glabrous or pubescent; limb 0.75–1.25 mm. long, dentate but with teeth usually less than half the length of the tube, ciliate. Corolla creamy white; tube 2.25–3 mm. long, with a ring of deflexed hairs set just below top inside; lobes lanceolate to ovate, 1.5–2.25 mm. long, 1–1.75 mm. wide, acute. Anthers fully exserted but seldom reflexing. Style 4–6.5 mm. long, glabrous; stigmatic knob 0.75–1.25 mm. long. Disk pubescent. Fruit black when mature, ± broadly oblong in outline, 0.8–1.1 cm. long, 1.1–1.5 cm. wide, slightly indented, glabrous; pyrene hemispherical to suborbicular, 6–7 mm. long, 5–6 mm. wide at apex; area from point of attachment lying perpendicular to ventral face triangular, containing a small crest.

Morphology

habit: climber

habit: shrub

Habitat

In evergreen forest understorey, swamp forest (mushitu) and riverine vegetation, in mixed deciduous woodlands on Kalahari Sand and in miombo woodlands often on termitaria, on sandy soils

HAB. Forest edges and scrub; 275–1525 m.

Distribution

Mozambique Maputo, corolla fallen 1908, Sim 20860 (PRE).

Kenya Shimba Hills, Godoni Forest area, 2–3 km. N. of Kwale Forest Station, 16–23 Sept. 1976, Spjut 4582! & near Giriama Point, 18 Sept. 1968, Gillett 18704! & 1927, Gardner 1435!

Uganda Serere, July 1932, Chandler 822!

Zimbabwe Chirinda Forest, fl. & fr. 18.x.1947, Wild 2026 (K; SRGH).

Zambia Ndola, fr. 8.xii.1953, Fanshawe 556 (K).

Uganda Koich R., near Rumogi, Eggeling I860!

Malawi Nkhata Bay Distr., border of Chombe Tea Estate near Mweza Village, fl. 13.iv.1960, Adlard 337 (K; LISC; SRGH).

Mozambique Sofala Prov., Inhansato, region of Inhaminga, fl. 13.iii.1955, Gomes e Sousa 4286 (COI; K).

Zambia Pemba, fl. 1933, Trapnell 1303 (K).

Uganda Sese [Sesse] Is., Bubembe I., 20 Jan. 1956, Dawkins 859!

Tanzania W. Usambara Mts., Bumbuli at DC’s rest-house, 11 May 1953, Drummond & Hemsley 2492!

Tanzania Rondo Plateau, Nandembo, 11 Dec. 1955, Milne-Redhead & Taylor 7638!

Zambia Kabompo Distr., 59 km WSW of Kabompo on Zambezi (Balovale) road, fl. bud 24.iii.1961, Drummond & Rutherford-Smith 7305 (K; LISC; PRE; SRGH).

Malawi Mulanje Distr., Likhubula River, above Likhubula House, west side of Mt. Mulanje, fl. 6.v.1980, Blackmore & Brummitt 1467 (K).

Zambia Chinsali Distr., near Great North Road, 48 km north of Mpika, fr. 28.xi.1952, White 3763 (FHO; K).

Mozambique Milange, fl. 24.ii.1943, Torre 4831 (LISC).

Tanzania Kungwe Mts., Kasiha [Kasieha] R., 20 July 1959, Harley & Newbould 4479!

Mozambique between Mocimboa da Praia and Diaca, c. 20 km from Mocimboa da Praia, fl. bud 25.iii.1961, Gomes e Sousa 4666 (COI; K; PRE).

Zambia Kamaila Forest Reserve, 40 km north of Lusaka, fr. 12.ix.1963, Angus 3735 (FHO; K); Luangwa Valley, Mutinondo Stream, Muchinga Escarpment, corolla fallen 23.viii.1966, Astle 4975 (K).

Literature

in Kew Bull. 41: 974 (1986). —K. Coates Palgrave, Trees Southern Africa, ed. 3, rev.: 16k of appendix (1988). —Bridson in F.T.E.A., Rubiaceae: 914 (1991). —Beentje, Kenya Trees, Shrubs Lianas: 518 (1994). Type from Uganda.

in K.B. 41: 974 (1986).

Distribution Map

 
  • Native distribution
Found in
  • Africa East Tropical Africa Kenya
  • Tanzania
  • Uganda
  • Northeast Tropical Africa Sudan
  • South Tropical Africa Angola
  • Malawi
  • Mozambique
  • Zambia
  • Zimbabwe
  • West Tropical Africa Benin
  • Burkina
  • Ghana
  • Guinea
  • Guinea-Bissau
  • Ivory Coast
  • Mali
  • Nigeria
  • Senegal
  • Sierra Leone
  • Togo
  • West-Central Tropical Africa Burundi
  • Cameroon
  • Central African Republic
  • Congo
  • Gabon
  • Rwanda
  • Zaire

  Bibliography

  • 1 Lejoy, J., Ndjele, M.-B. & Geerinck, D. (2010). Catalogue-flore des plantes vasculaires des districts de Kisangani et de la Tshopo (RD Congo). Taxonomania. Revue de Taxonomie et de Nomenclature Botaniques 30: 1-307.
  • 2 Timberlake, J.R., Bayliss, J., Alves, T., Francisco, J., Harris, T., Nangoma, D. & de Sousa, C. (2009). Biodiversity and Conservation of Mchese Mountain, Malawi. Report produced under the Darwin Initiative Award 15/036: 1-71. Royal Botanic Gardens, Kew.
  • 3 Figueiredo, E. & Smith, G.F. (2008). Plants of Angola. Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
  • 4 Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
  • 5 Sosef, M.S.M. & al. (2006). Check-list des plantes vasculaires du Gabon. Scripta Botanica Belgica 35: 1-438.
  • 6 Strugnell, A.M. (2006). A checklist of the Spermatophytes of Mt. Mulanje, Malawi. Scripta Botanica Belgica 34: 1-199.
  • 7 Ruhsam, M (2005). Survey of all the Rubiaceae specimens at the Royal Botanic Gardens, Kew.
  • 8 Sita, P. & Moutsambote, J.-M. (2005). Catalogue des plantes vasculaires du Congo , ed. sept. 2005: 1-158. ORSTOM, Centre de Brazzaville.
  • 9 Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.
  • 10 Harris, D.J. (2002). The vascular plants of the Dzanga-Sangha Reserve, Central African Republic: 1-274. National Botanic Garden (Belgium), Meise.
  • 11 B. Verdcourt & D. Bridson (1991) Rubiaceae. Flora of Tropical East Africa 0 (0)
  • 12 Brunel, J.F., Hiepo, P. & Scholz, H. (eds.) (1984). Flore Analytique du Togo Phanérogames: 1-751. GTZ, Eschborn.

 Information From

Flora Zambesiaca
http://apps.kew.org/efloras/
Flora Zambesiaca. Published on the Internet; http://apps.kew.org/efloras/
Flora of Tropical East Africa
http://apps.kew.org/efloras/
Flora of Tropical East Africa. Published on the Internet; http://apps.kew.org/efloras/
WCS higher taxonomy
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World Checklist of Selected Plant Families
http://apps.kew.org/wcsp/
WCSP 2014. 'World Checklist of Selected Plant Families. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet; http://apps.kew.org/wcsp/ Retrieved 2011 onwards
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